The example will be calculated for the gene-species which is selected in the left Hitlist of the 'TRANSFER SPECIES' window.
The ACI expression applies to the Organism or the Gene if 'Copy from organism/gene' is selected. If 'Only use ACI' is selected it applies to the gene-species.
Note that it is possible to create hierarchical keys. Simply enter something like 'organism/full_name' into the destination field.